Two new species of euptychia hübner, 1818 from the upper Amazon basin (Lepidoptera, Nymphalidae, Satyrinae)

Andrew F.E. Neild, Shinichi Nakahara, Thamara Zacca, Steven Fratello, Gerardo Lamas, Jean François Le Crom, Diego R. Dolibaina, Fernando M.S. Dias, Mirna M. Casagrande, Olaf H.H. Mielke, Marianne Espeland

Research output: Contribution to journalArticlepeer-review

7 Scopus citations


E. Neild et al. Two new species of Euptychia Hübner, 1818 are described from the upper Amazon basin: E. attenboroughi Neild, Nakahara, Fratello & Le Crom, sp. n. (type locality: Amazonas, Venezuela), and E. sophiae Zacca, Nakahara, Dolibaina & Dias, sp. n. (type locality: Acre, Brazil). Their unusual facies prompted molecular and phylogenetic analyses of one of the species resulting in support for their classification in monophyletic Euptychia. Diagnostic characters for the two species are presented based on wing morphology, wing pattern, presence of androconial patches on the hindwing, and genitalia. Our results indicate that the projection of the tegumen above the uncus, previously considered a synapomorphy for Euptychia, is not shared by all species in the genus. The adults and their genitalia are documented, and distribution data and a map are provided.
Original languageAmerican English
Pages (from-to)87-108
Number of pages22
Issue number541
StatePublished - 1 Jan 2015


  • Brazil
  • Colombia
  • Euptychiina
  • Mitochondrial DNA COI barcode
  • Neotropics
  • Peru
  • South America
  • Venezuela


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